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Back in the September release, Marketo changed the URLs in emails to be shorter. We notified everyone that the old, longer, links will expire 90 days after the release. I believe we just passed the 90 day mark and that is why your links are no longer working.
More info here: https://community.marketo.com/MarketoArticle?id=kA050000000LBSsCAO#shorter-urls
Ah-ha! Thanks, Will. I forgot that that was going to happen.