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When a lead is synced to Salesforce via the campaign sync rather than a 'Sync lead to SFDC' flow step, Marketo refers to the lead assignment rules that are set up inside of Salesforce.
If you are an admin in Salesforce, go to Setup > Leads > Lead Assignment rules to see whether any assignment rules are set up for your organisation.
If no assignment rules are set up, the new lead will be assigned to the Default Lead Owner (Setup > Leads > Lead Settings)
Hope this helps!
Thanks Suzanne! Just what I was looking for.